Full
model name: Hasegawa et.al., 1985 with 2 rate variation parameters drawn from
two independent unit gamma distributions.
Model type:
Nucleotide
Model
options:
The user selects how many classes transition rates and
transversion rates shall have.
Independent
Parameters:
a -
branch length, R - global transversion/transition rate,
alpha - shape parameter of the unit gamma distribution
of transversion rates, beta - shape parameter of the
unit gamma distribution of transition rates.
Rate Matrix:
with the vector representing
nucleotide frequencies (proportions) from the data.
is distributed as unit
gamma with shape parameter alpha, and
is distributed as unit gamma with shape parameter beta.
'*' is defined to be the negative of the sum of all other row elements.
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